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On spellcaster roles like Priest, definitely wait until about Level 14, or until you get all the Vine spells you want (including the high level spells, especially Breath of Air). Same with Wizards: make sure you have all the Stone spells and Sun spells you want, but especially Stone spells, as the only other roles that can learn them are Zenmaster and Valkyrie near the endgame.
On Warriors and Rogues, you can ascend them ASAP.
-Sazerac
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