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In all the games so far in which I did the Alliance thingy, I have always given the T'rang letter to the Mook before even doing any Umpani quests after the obstacle course (in other words, giving the Mook the T'rang letter without having the Umpani letter in my inv), and I have never once had Z'ant go hostile on me (knock on wood).
So as far as I can tell, you just might be on to something.
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