One more discovery I made on my third time through: this time I went for the heart as soon as I entered the area, and it wasn't there! Then I defeated both guardian demons, got both keys and went back. Then it was there. I think you need to defeat the one in the red area, and then jump down the pit in his room. That will splash you to the wall close to the elevator, which I think is the hint to find the heart.
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